Themeda villosa

(Poir.) A.Camus (1922)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Poales Family: Poaceae Genus: Themeda

lyon's grass

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Description

Key Characters:

Growth Form: Robust perennials.

Stems: Culms tufted, stout, 20–40 dm tall, solid, culm base compressed flabellate.

Roots: Fibrous root system.

Leaves: Leaves simple. Alternate. Blades up to 150 cm long, up to 20 mm wide. Apex acuminate. Base narrowed to the sheath. Surfaces glabrous, midrib white, prominent, margins and nerves scabrous on upper surface. Margins entire. Veins parallel. Sheaths keeled, ± compressed, margins overlapping, glabrous; ligule membranous, ciliate. Stipules absent.

Flowers: Flowers in inflorescences large compound, drooping, paniculate, spathes 3–3.5 cm long, subtending a flattened rachis, apex pubescent, racemes of spikelet pairs at about the middle of the spathe. involucral and pedicellate spikelets reddish brown, similar in shape, narrowly fusiform, acuminate, 15–20 mm long, pedicels rufous pubescent; sessile spikelet with a callus ca. 2 mm long, densely pubescent with rufous hairs, first glume ovate, 10–nerved, the nerves reticulate toward apex, densely rufous pubescent between the nerves, apex acute, first lemma 1–nerved, apex acute; second lemma 1–nerved, hyaline, prolonged into a twisted pubescent awn up to 42 mm long, second palea lanceolate, nerveless, hyaline. Stamens 3 (when present). Ovary superior.

Fruit: Caryopsis reddish brown; fusiform; ca. 5 mm long.

Ploidy: 2n = 32; 40

Habitat: Dry pastures; along roadsides; and other disturbed sites.

Elevation Range: 0–700 m.

Historical Distribution

Images

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Bibliography

Name Published In: H.Lecomte, Fl. Indo-Chine 7: 364 (1922)

Other References

Wagner et al. 1990:1599 (K, O, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 decaying leaves on forest floor PRESERVED_SPECIMEN baker, r.f. collector number: 61 Maui BISH 4/9/1975
2 Themeda villosa honolulu; fair grounds Small trees at edge of forest. PRESERVED_SPECIMEN lee, h.a. weller collector number: 121 Oahu BISH 12/15/1924
3 Themeda villosa Hanamaulu, Lihue, near beach In pasture. PRESERVED_SPECIMEN Hosaka, E.Y. 1655 Kauai BISH 11/4/1936
4 Themeda villosa kawailoa training area, drum road, kamananui gulch Shaded 10 ft. deep depression of lava tube opening, growing with Pipturus albidus, Physalis peruviana, Bryophyllum pinnatum, the ferns Nephrolepis multiflora and Adiantum capillus-veneris. PRESERVED_SPECIMEN kawelo, k. collector number: s.n. Maui BISH 6/4/2003
5 Themeda villosa Koolau Mountains; Moanalua Valley Common roadside weed. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
6 Themeda villosa laupahoehoe section of hilo forest reserve, north hilo district Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN cuddihy, l.w. collector number: 923 Hawaii BISH 11/2/1981
7 Themeda villosa koloa district. wahiawa, along dirt road from alexander's reservoir to kanaele bog. wet bank of gully PRESERVED_SPECIMEN wood, k.r. nishek, r.; clark, m. collector number: 14310 Maui BISH 9/10/2010
8 Themeda villosa kona, keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
9 Themeda villosa Junction of Highways 18 & 19, above Kailua-Kona PRESERVED_SPECIMEN Herbst, D.R. 5061 Hawaii BISH 10/8/1974
10 Themeda villosa kona; keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
11 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. Palmer, R. Collector Number: 3748 Hawaii BISH 1/14/1998
12 Themeda villosa Koolau mountains; Moanalua Valley, Damon Estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
13 Themeda villosa Kealakekua PRESERVED_SPECIMEN Toy, B. s.n. Hawaii BISH 5/1/1979
14 Themeda villosa Honolulu; Lyon Arboretum, on makai side of access road ca. 200 yards from seismograph station. Below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN Baker, R.F. 61 Oahu BISH 4/9/1975
15 Themeda villosa Koloa District. Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha-Dicranopteris linearis wet forest. Antidesma platyphylla, Ilex anomala, Polyscias waialeale, Scaevola gaudichaudiana, Bobea brevipes, Psychotria mariniana, weeds: Psidum cattleianum PRESERVED_SPECIMEN Perlman, S. 23477 Kauai BISH 5/23/2013
16 Themeda villosa Kona, before descent to Kailua Wet area. PRESERVED_SPECIMEN Christensen, C. 2 Hawaii BISH 11/14/1963
17 Themeda villosa hanamaulu, lihue, near beach Occasional in near beach in dry, rocky pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Lanai BISH 11/4/1936
18 Themeda villosa Along highway between Keaau and Pahoa. PRESERVED_SPECIMEN Nagata, K.M. 4110 Hawaii BISH 5/14/1990
19 Themeda villosa Koloa District. Wahiawa, along dirt road from Alexander's reservoir to Kanaele bog. Secondary invasive vegetatino. W/ native Metrosideros polymrpha and Dicranopteris linearis. PRESERVED_SPECIMEN Wood, K.R. 14310 Kauai BISH 9/10/2010
20 Themeda villosa Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua. Acacia koa-Metrosideros sp. lowland mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
21 Themeda villosa Hilo Forest Reserve, Saddle Rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 NY 7/7/1997
22 Themeda villosa hilo forest reserve, saddle rd at 21.7km Roadside PRESERVED_SPECIMEN c. r. annable 3748 NY 1/14/1998
23 Themeda villosa Hilo Forest Reserve, Saddle Rd at 21.7km Roadside PRESERVED_SPECIMEN C. R. Annable 3748 NY 1/14/1998
24 Themeda villosa koloa district wahiawa PRESERVED_SPECIMEN k. r. wood 14310 NY 9/10/2010
25 Themeda villosa hilo forest reserve, saddle rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN c. r. annable 3604 NY 7/7/1997
26 Themeda villosa Lyon Arboretum. On makai side of access road about 200' down road from seismograph station. Below section 31. vSunny. PRESERVED_SPECIMEN R. F. Baker 61 Oahu PTBG 4/9/1975
27 Themeda villosa OAHU: Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua; Acacia koa-Metrosideros sp. lowland mesic to wet forest w/ Psychotria mariniana, Syzygium sandwicensis, Scaevola chamissoniana. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
28 Themeda villosa Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha- Dicranopteris linearis wet forest PRESERVED_SPECIMEN Steve Perlman 23477 Kauai PTBG 5/23/2013
29 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve. Growing beside jeep road. Logged ohia-koa forest. PRESERVED_SPECIMEN L. W. Cuddihy 923 Hawaii PTBG 11/2/1981
30 Themeda villosa honolulu PRESERVED_SPECIMEN h. lee 121 Oahu US 8/8/1924
31 Themeda villosa Wahiawa, along dirt road from Alexander's Reservoir to Kanaele Bog. Secondary invasive vegetation. PRESERVED_SPECIMEN K. R. Wood 14310 Kauai PTBG 9/10/2010
32 Themeda villosa Kona, Keauhou. PRESERVED_SPECIMEN A. F. Judd s.n. Hawaii PTBG 11/27/1937
33 Themeda villosa Haleakala Substation, Makawao PRESERVED_SPECIMEN Hosaka, E. Y. 2443 Maui US 1939-00-00
34 Themeda villosa Wahiawa Mountains. Growing along jeep track accessing Kanaele Bog above Kalaheo. PRESERVED_SPECIMEN Clay Trauernicht 594 Kauai PTBG 10/26/2008
35 Themeda villosa Waipio Valley PRESERVED_SPECIMEN Degener, O.; Degener, I. 29849 Oahu US 1/20/1963
36 Themeda villosa waipio valley PRESERVED_SPECIMEN o. degener & i. degener 29849 Oahu US 1/20/1963
37 Themeda villosa Honolulu PRESERVED_SPECIMEN Lee, H. A. 121 Oahu US 4/7/1905
38 Themeda villosa jct of hwy 18 and 19, above kailua-kona. PRESERVED_SPECIMEN d. r. herbst & s. ishikawa 5061 Hawaii US 10/7/1974
39 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii US 7/7/1997
40 Themeda villosa hilo forest reserve, 16mi marker on saddle rd PRESERVED_SPECIMEN c. r. annable, r. palmer, d. paul & d. ward 3604 Hawaii US 7/7/1997
41 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN Annable, C. R.; Palmer, R.; Paul, D.; Ward, D. 3604 Hawaii US 7/7/1997
42 Themeda villosa Hanamaulu, Lihue. In pasture near beach. PRESERVED_SPECIMEN E. Y. Hosaka 1655 Kauai PTBG 11/4/1936
43 Themeda villosa kealakekua PRESERVED_SPECIMEN toy, b. collector number: s.n. Maui BISH 5/1/1979
44 Themeda villosa Jct of Hwy 18 and 19, above Kailua-Kona. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 5061 Hawaii US 10/7/1974
45 Themeda villosa south hilo distr., scenic drive along hamakua coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN staples, g.w. purell, m. collector number: 1268 Hawaii BISH 2/13/2007
46 Themeda villosa sw of dillingham ranch, mokuleia PRESERVED_SPECIMEN o. degener, d. l. topping, -- martinez & -- salucop 11058 Oahu US 3/12/1937
47 Themeda villosa Honolulu; Fair grounds PRESERVED_SPECIMEN Lee, H.A. 121 Oahu BISH 12/15/1924
48 Themeda villosa jeep road between wahiawa bog (kanaele swamp) and trail to tv relay tower on mt. kahili PRESERVED_SPECIMEN wagner, w.l. mill, s.; flynn,t.; peterson, b.; peterson, g. collector number: 5199 Kauai BISH 12/22/1983
49 Themeda villosa haleakala substation, makawao PRESERVED_SPECIMEN e. y. hosaka 2443 Maui US 4/9/1939
50 Themeda villosa SW of Dillingham Ranch, Mokuleia PRESERVED_SPECIMEN Degener, O.; Topping, D. L.; Martinez, --; Salucop, -- 11058 Oahu US 3/12/1937
51 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. Collector Number: 923 Hawaii BISH 11/2/1981
52 Themeda villosa hilo forest reserve; saddle rd. at 21.7 km ridge; with Holcus, Acacia, Hypochoeris, Styphelia PRESERVED_SPECIMEN annable, c.r. palmer, r. collector number: 3748 Maui BISH 1/14/1998
53 Themeda villosa hanamaulu, lihue, near beach PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
54 Themeda villosa Tree Planting Road near Stainback highway Ruderal. PRESERVED_SPECIMEN Stemmermann, R.L. 6703 Hawaii BISH 1/6/1981
55 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. 3748 Hawaii BISH 1/14/1998
56 Themeda villosa Kona; Keauhou PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 11/12/1937
57 Themeda villosa Kona Along roadside. PRESERVED_SPECIMEN Hosaka, E.Y. s.n. Hawaii BISH 9/1/1936
58 Themeda villosa Jeep road between Wahiawa Bog (Kanaele Swamp) and trail to TV relay tower on Mt. Kahili PRESERVED_SPECIMEN Wagner, W.L. 5199 Kauai BISH 12/22/1983
59 Themeda villosa Kawailoa Training Area, Drum Road, Kamananui Gulch Roadside vegetation, along with Hyparrhenia rufa, Dicranopteris linearis, Panicum maximum. PRESERVED_SPECIMEN Kawelo, K. s.n. Oahu BISH 6/4/2003
60 Themeda villosa koolau mountains; moanalua valley, damon estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN herbst, d.r. collector number: 355 Oahu BISH 3/18/1967
61 Themeda villosa Moanalua Valley, Campsite No. 1 Common along stream and jeep trail up to about 800 ft. PRESERVED_SPECIMEN Herbst, D.R. 1523 Oahu BISH 1/15/1970
62 Themeda villosa tree planting road near stainback highway Ruderal. PRESERVED_SPECIMEN stemmermann, r.l. collector number: 6703 Hawaii BISH 1/6/1981
63 Themeda villosa junction of highways 18 & 19, above kailua-kona PRESERVED_SPECIMEN herbst, d.r. ishikawa, s. collector number: 5061 Hawaii BISH 10/7/1974
64 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. 923 Hawaii BISH 11/2/1981
65 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN baker, r.f. collector number: 61 Oahu BISH 4/9/1975
66 Themeda villosa hanamaulu, lihue, near beach In pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
67 Themeda villosa South Hilo distr., Scenic Drive along Hamakua Coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN Staples, G. 1268 Hawaii BISH 2/13/2007
68 Themeda villosa Honokohau Valley, Kava Kamp Roadside PRESERVED_SPECIMEN C. R. Annable 3748 Maui HAPI 1/14/1998
69 Themeda villosa Oahu Islands, Waianae Range, Waikele Gulch Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 Oahu HAPI 7/7/1997
70 Themeda villosa southwest of dillingham ranch Single plant in pasture. PRESERVED_SPECIMEN degener, o. topping, d.l.; martinez, m.; salucop, f. collector number: 11058 Oahu BISH 3/12/1937
71 Themeda villosa Southwest of Dillingham Ranch Single plant in pasture. PRESERVED_SPECIMEN Degener, O. 11058 Oahu BISH 3/12/1937
72 Themeda villosa kona PRESERVED_SPECIMEN hosaka, e.y. collector number: s.n. Oahu BISH 9/1/1936
73 Themeda villosa kona, before descent to kailua Wet area. PRESERVED_SPECIMEN christensen, c. collector number: 2 Hawaii BISH 11/14/1963
74 Themeda villosa koolau mountains; moanalua valley PRESERVED_SPECIMEN herbst, d.r. collector number: 355 BISH 3/18/1967
75 Themeda villosa Hoea PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii HAPI 7/7/1997
76 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 decaying leaves on forest floor PRESERVED_SPECIMEN baker, r.f. collector number: 61 Maui BISH 4/9/1975
77 Themeda villosa honolulu; fair grounds Small trees at edge of forest. PRESERVED_SPECIMEN lee, h.a. weller collector number: 121 Oahu BISH 12/15/1924
78 Themeda villosa Hanamaulu, Lihue, near beach In pasture. PRESERVED_SPECIMEN Hosaka, E.Y. 1655 Kauai BISH 11/4/1936
79 Themeda villosa kawailoa training area, drum road, kamananui gulch Shaded 10 ft. deep depression of lava tube opening, growing with Pipturus albidus, Physalis peruviana, Bryophyllum pinnatum, the ferns Nephrolepis multiflora and Adiantum capillus-veneris. PRESERVED_SPECIMEN kawelo, k. collector number: s.n. Maui BISH 6/4/2003
80 Themeda villosa Koolau Mountains; Moanalua Valley Common roadside weed. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
81 Themeda villosa laupahoehoe section of hilo forest reserve, north hilo district Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN cuddihy, l.w. collector number: 923 Hawaii BISH 11/2/1981
82 Themeda villosa koloa district. wahiawa, along dirt road from alexander's reservoir to kanaele bog. wet bank of gully PRESERVED_SPECIMEN wood, k.r. nishek, r.; clark, m. collector number: 14310 Maui BISH 9/10/2010
83 Themeda villosa kona, keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
84 Themeda villosa Junction of Highways 18 & 19, above Kailua-Kona PRESERVED_SPECIMEN Herbst, D.R. 5061 Hawaii BISH 10/8/1974
85 Themeda villosa kona; keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
86 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. Palmer, R. Collector Number: 3748 Hawaii BISH 1/14/1998
87 Themeda villosa Koolau mountains; Moanalua Valley, Damon Estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
88 Themeda villosa Kealakekua PRESERVED_SPECIMEN Toy, B. s.n. Hawaii BISH 5/1/1979
89 Themeda villosa Honolulu; Lyon Arboretum, on makai side of access road ca. 200 yards from seismograph station. Below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN Baker, R.F. 61 Oahu BISH 4/9/1975
90 Themeda villosa Koloa District. Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha-Dicranopteris linearis wet forest. Antidesma platyphylla, Ilex anomala, Polyscias waialeale, Scaevola gaudichaudiana, Bobea brevipes, Psychotria mariniana, weeds: Psidum cattleianum PRESERVED_SPECIMEN Perlman, S. 23477 Kauai BISH 5/23/2013
91 Themeda villosa Kona, before descent to Kailua Wet area. PRESERVED_SPECIMEN Christensen, C. 2 Hawaii BISH 11/14/1963
92 Themeda villosa hanamaulu, lihue, near beach Occasional in near beach in dry, rocky pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Lanai BISH 11/4/1936
93 Themeda villosa Along highway between Keaau and Pahoa. PRESERVED_SPECIMEN Nagata, K.M. 4110 Hawaii BISH 5/14/1990
94 Themeda villosa Koloa District. Wahiawa, along dirt road from Alexander's reservoir to Kanaele bog. Secondary invasive vegetatino. W/ native Metrosideros polymrpha and Dicranopteris linearis. PRESERVED_SPECIMEN Wood, K.R. 14310 Kauai BISH 9/10/2010
95 Themeda villosa Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua. Acacia koa-Metrosideros sp. lowland mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
96 Themeda villosa Hilo Forest Reserve, Saddle Rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 NY 7/7/1997
97 Themeda villosa hilo forest reserve, saddle rd at 21.7km Roadside PRESERVED_SPECIMEN c. r. annable 3748 NY 1/14/1998
98 Themeda villosa Hilo Forest Reserve, Saddle Rd at 21.7km Roadside PRESERVED_SPECIMEN C. R. Annable 3748 NY 1/14/1998
99 Themeda villosa koloa district wahiawa PRESERVED_SPECIMEN k. r. wood 14310 NY 9/10/2010
100 Themeda villosa hilo forest reserve, saddle rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN c. r. annable 3604 NY 7/7/1997
101 Themeda villosa Lyon Arboretum. On makai side of access road about 200' down road from seismograph station. Below section 31. vSunny. PRESERVED_SPECIMEN R. F. Baker 61 Oahu PTBG 4/9/1975
102 Themeda villosa OAHU: Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua; Acacia koa-Metrosideros sp. lowland mesic to wet forest w/ Psychotria mariniana, Syzygium sandwicensis, Scaevola chamissoniana. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
103 Themeda villosa Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha- Dicranopteris linearis wet forest PRESERVED_SPECIMEN Steve Perlman 23477 Kauai PTBG 5/23/2013
104 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve. Growing beside jeep road. Logged ohia-koa forest. PRESERVED_SPECIMEN L. W. Cuddihy 923 Hawaii PTBG 11/2/1981
105 Themeda villosa honolulu PRESERVED_SPECIMEN h. lee 121 Oahu US 8/8/1924
106 Themeda villosa Wahiawa, along dirt road from Alexander's Reservoir to Kanaele Bog. Secondary invasive vegetation. PRESERVED_SPECIMEN K. R. Wood 14310 Kauai PTBG 9/10/2010
107 Themeda villosa Kona, Keauhou. PRESERVED_SPECIMEN A. F. Judd s.n. Hawaii PTBG 11/27/1937
108 Themeda villosa Haleakala Substation, Makawao PRESERVED_SPECIMEN Hosaka, E. Y. 2443 Maui US 1939-00-00
109 Themeda villosa Wahiawa Mountains. Growing along jeep track accessing Kanaele Bog above Kalaheo. PRESERVED_SPECIMEN Clay Trauernicht 594 Kauai PTBG 10/26/2008
110 Themeda villosa Waipio Valley PRESERVED_SPECIMEN Degener, O.; Degener, I. 29849 Oahu US 1/20/1963
111 Themeda villosa waipio valley PRESERVED_SPECIMEN o. degener & i. degener 29849 Oahu US 1/20/1963
112 Themeda villosa Honolulu PRESERVED_SPECIMEN Lee, H. A. 121 Oahu US 4/7/1905
113 Themeda villosa jct of hwy 18 and 19, above kailua-kona. PRESERVED_SPECIMEN d. r. herbst & s. ishikawa 5061 Hawaii US 10/7/1974
114 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii US 7/7/1997
115 Themeda villosa hilo forest reserve, 16mi marker on saddle rd PRESERVED_SPECIMEN c. r. annable, r. palmer, d. paul & d. ward 3604 Hawaii US 7/7/1997
116 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN Annable, C. R.; Palmer, R.; Paul, D.; Ward, D. 3604 Hawaii US 7/7/1997
117 Themeda villosa Hanamaulu, Lihue. In pasture near beach. PRESERVED_SPECIMEN E. Y. Hosaka 1655 Kauai PTBG 11/4/1936
118 Themeda villosa kealakekua PRESERVED_SPECIMEN toy, b. collector number: s.n. Maui BISH 5/1/1979
119 Themeda villosa Jct of Hwy 18 and 19, above Kailua-Kona. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 5061 Hawaii US 10/7/1974
120 Themeda villosa south hilo distr., scenic drive along hamakua coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN staples, g.w. purell, m. collector number: 1268 Hawaii BISH 2/13/2007
121 Themeda villosa sw of dillingham ranch, mokuleia PRESERVED_SPECIMEN o. degener, d. l. topping, -- martinez & -- salucop 11058 Oahu US 3/12/1937
122 Themeda villosa Honolulu; Fair grounds PRESERVED_SPECIMEN Lee, H.A. 121 Oahu BISH 12/15/1924
123 Themeda villosa jeep road between wahiawa bog (kanaele swamp) and trail to tv relay tower on mt. kahili PRESERVED_SPECIMEN wagner, w.l. mill, s.; flynn,t.; peterson, b.; peterson, g. collector number: 5199 Kauai BISH 12/22/1983
124 Themeda villosa haleakala substation, makawao PRESERVED_SPECIMEN e. y. hosaka 2443 Maui US 4/9/1939
125 Themeda villosa SW of Dillingham Ranch, Mokuleia PRESERVED_SPECIMEN Degener, O.; Topping, D. L.; Martinez, --; Salucop, -- 11058 Oahu US 3/12/1937
126 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. Collector Number: 923 Hawaii BISH 11/2/1981
127 Themeda villosa hilo forest reserve; saddle rd. at 21.7 km ridge; with Holcus, Acacia, Hypochoeris, Styphelia PRESERVED_SPECIMEN annable, c.r. palmer, r. collector number: 3748 Maui BISH 1/14/1998
128 Themeda villosa hanamaulu, lihue, near beach PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
129 Themeda villosa Tree Planting Road near Stainback highway Ruderal. PRESERVED_SPECIMEN Stemmermann, R.L. 6703 Hawaii BISH 1/6/1981
130 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. 3748 Hawaii BISH 1/14/1998
131 Themeda villosa Kona; Keauhou PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 11/12/1937
132 Themeda villosa Kona Along roadside. PRESERVED_SPECIMEN Hosaka, E.Y. s.n. Hawaii BISH 9/1/1936
133 Themeda villosa Jeep road between Wahiawa Bog (Kanaele Swamp) and trail to TV relay tower on Mt. Kahili PRESERVED_SPECIMEN Wagner, W.L. 5199 Kauai BISH 12/22/1983
134 Themeda villosa Kawailoa Training Area, Drum Road, Kamananui Gulch Roadside vegetation, along with Hyparrhenia rufa, Dicranopteris linearis, Panicum maximum. PRESERVED_SPECIMEN Kawelo, K. s.n. Oahu BISH 6/4/2003
135 Themeda villosa koolau mountains; moanalua valley, damon estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN herbst, d.r. collector number: 355 Oahu BISH 3/18/1967
136 Themeda villosa Moanalua Valley, Campsite No. 1 Common along stream and jeep trail up to about 800 ft. PRESERVED_SPECIMEN Herbst, D.R. 1523 Oahu BISH 1/15/1970
137 Themeda villosa tree planting road near stainback highway Ruderal. PRESERVED_SPECIMEN stemmermann, r.l. collector number: 6703 Hawaii BISH 1/6/1981
138 Themeda villosa junction of highways 18 & 19, above kailua-kona PRESERVED_SPECIMEN herbst, d.r. ishikawa, s. collector number: 5061 Hawaii BISH 10/7/1974
139 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. 923 Hawaii BISH 11/2/1981
140 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN baker, r.f. collector number: 61 Oahu BISH 4/9/1975
141 Themeda villosa hanamaulu, lihue, near beach In pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
142 Themeda villosa South Hilo distr., Scenic Drive along Hamakua Coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN Staples, G. 1268 Hawaii BISH 2/13/2007
143 Themeda villosa Honokohau Valley, Kava Kamp Roadside PRESERVED_SPECIMEN C. R. Annable 3748 Maui HAPI 1/14/1998
144 Themeda villosa Oahu Islands, Waianae Range, Waikele Gulch Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 Oahu HAPI 7/7/1997
145 Themeda villosa southwest of dillingham ranch Single plant in pasture. PRESERVED_SPECIMEN degener, o. topping, d.l.; martinez, m.; salucop, f. collector number: 11058 Oahu BISH 3/12/1937
146 Themeda villosa Southwest of Dillingham Ranch Single plant in pasture. PRESERVED_SPECIMEN Degener, O. 11058 Oahu BISH 3/12/1937
147 Themeda villosa kona PRESERVED_SPECIMEN hosaka, e.y. collector number: s.n. Oahu BISH 9/1/1936
148 Themeda villosa kona, before descent to kailua Wet area. PRESERVED_SPECIMEN christensen, c. collector number: 2 Hawaii BISH 11/14/1963
149 Themeda villosa koolau mountains; moanalua valley PRESERVED_SPECIMEN herbst, d.r. collector number: 355 BISH 3/18/1967
150 Themeda villosa Hoea PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii HAPI 7/7/1997
151 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 decaying leaves on forest floor PRESERVED_SPECIMEN baker, r.f. collector number: 61 Maui BISH 4/9/1975
152 Themeda villosa honolulu; fair grounds Small trees at edge of forest. PRESERVED_SPECIMEN lee, h.a. weller collector number: 121 Oahu BISH 12/15/1924
153 Themeda villosa Hanamaulu, Lihue, near beach In pasture. PRESERVED_SPECIMEN Hosaka, E.Y. 1655 Kauai BISH 11/4/1936
154 Themeda villosa kawailoa training area, drum road, kamananui gulch Shaded 10 ft. deep depression of lava tube opening, growing with Pipturus albidus, Physalis peruviana, Bryophyllum pinnatum, the ferns Nephrolepis multiflora and Adiantum capillus-veneris. PRESERVED_SPECIMEN kawelo, k. collector number: s.n. Maui BISH 6/4/2003
155 Themeda villosa Koolau Mountains; Moanalua Valley Common roadside weed. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
156 Themeda villosa laupahoehoe section of hilo forest reserve, north hilo district Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN cuddihy, l.w. collector number: 923 Hawaii BISH 11/2/1981
157 Themeda villosa koloa district. wahiawa, along dirt road from alexander's reservoir to kanaele bog. wet bank of gully PRESERVED_SPECIMEN wood, k.r. nishek, r.; clark, m. collector number: 14310 Maui BISH 9/10/2010
158 Themeda villosa kona, keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
159 Themeda villosa Junction of Highways 18 & 19, above Kailua-Kona PRESERVED_SPECIMEN Herbst, D.R. 5061 Hawaii BISH 10/8/1974
160 Themeda villosa kona; keauhou PRESERVED_SPECIMEN judd jr., a.f. collector number: s.n. Hawaii BISH 11/12/1937
161 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. Palmer, R. Collector Number: 3748 Hawaii BISH 1/14/1998
162 Themeda villosa Koolau mountains; Moanalua Valley, Damon Estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN Herbst, D.R. 355 Oahu BISH 3/18/1967
163 Themeda villosa Kealakekua PRESERVED_SPECIMEN Toy, B. s.n. Hawaii BISH 5/1/1979
164 Themeda villosa Honolulu; Lyon Arboretum, on makai side of access road ca. 200 yards from seismograph station. Below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN Baker, R.F. 61 Oahu BISH 4/9/1975
165 Themeda villosa Koloa District. Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha-Dicranopteris linearis wet forest. Antidesma platyphylla, Ilex anomala, Polyscias waialeale, Scaevola gaudichaudiana, Bobea brevipes, Psychotria mariniana, weeds: Psidum cattleianum PRESERVED_SPECIMEN Perlman, S. 23477 Kauai BISH 5/23/2013
166 Themeda villosa Kona, before descent to Kailua Wet area. PRESERVED_SPECIMEN Christensen, C. 2 Hawaii BISH 11/14/1963
167 Themeda villosa hanamaulu, lihue, near beach Occasional in near beach in dry, rocky pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Lanai BISH 11/4/1936
168 Themeda villosa Along highway between Keaau and Pahoa. PRESERVED_SPECIMEN Nagata, K.M. 4110 Hawaii BISH 5/14/1990
169 Themeda villosa Koloa District. Wahiawa, along dirt road from Alexander's reservoir to Kanaele bog. Secondary invasive vegetatino. W/ native Metrosideros polymrpha and Dicranopteris linearis. PRESERVED_SPECIMEN Wood, K.R. 14310 Kauai BISH 9/10/2010
170 Themeda villosa Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua. Acacia koa-Metrosideros sp. lowland mesic to wet forest. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
171 Themeda villosa Hilo Forest Reserve, Saddle Rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 NY 7/7/1997
172 Themeda villosa hilo forest reserve, saddle rd at 21.7km Roadside PRESERVED_SPECIMEN c. r. annable 3748 NY 1/14/1998
173 Themeda villosa Hilo Forest Reserve, Saddle Rd at 21.7km Roadside PRESERVED_SPECIMEN C. R. Annable 3748 NY 1/14/1998
174 Themeda villosa koloa district wahiawa PRESERVED_SPECIMEN k. r. wood 14310 NY 9/10/2010
175 Themeda villosa hilo forest reserve, saddle rd at 27.6km Disturbed roadside in wet lowland PRESERVED_SPECIMEN c. r. annable 3604 NY 7/7/1997
176 Themeda villosa Lyon Arboretum. On makai side of access road about 200' down road from seismograph station. Below section 31. vSunny. PRESERVED_SPECIMEN R. F. Baker 61 Oahu PTBG 4/9/1975
177 Themeda villosa OAHU: Koolau Mts; Schofield-Waikane; drainage below trailhead; North Kaukonahua; Acacia koa-Metrosideros sp. lowland mesic to wet forest w/ Psychotria mariniana, Syzygium sandwicensis, Scaevola chamissoniana. PRESERVED_SPECIMEN K. R. Wood 5898 Oahu PTBG 1/2/1997
178 Themeda villosa Wahiawa area, along road to Kanaele bog from Alexander dam Metrosideros polymorpha- Dicranopteris linearis wet forest PRESERVED_SPECIMEN Steve Perlman 23477 Kauai PTBG 5/23/2013
179 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve. Growing beside jeep road. Logged ohia-koa forest. PRESERVED_SPECIMEN L. W. Cuddihy 923 Hawaii PTBG 11/2/1981
180 Themeda villosa honolulu PRESERVED_SPECIMEN h. lee 121 Oahu US 8/8/1924
181 Themeda villosa Wahiawa, along dirt road from Alexander's Reservoir to Kanaele Bog. Secondary invasive vegetation. PRESERVED_SPECIMEN K. R. Wood 14310 Kauai PTBG 9/10/2010
182 Themeda villosa Kona, Keauhou. PRESERVED_SPECIMEN A. F. Judd s.n. Hawaii PTBG 11/27/1937
183 Themeda villosa Haleakala Substation, Makawao PRESERVED_SPECIMEN Hosaka, E. Y. 2443 Maui US 1939-00-00
184 Themeda villosa Wahiawa Mountains. Growing along jeep track accessing Kanaele Bog above Kalaheo. PRESERVED_SPECIMEN Clay Trauernicht 594 Kauai PTBG 10/26/2008
185 Themeda villosa Waipio Valley PRESERVED_SPECIMEN Degener, O.; Degener, I. 29849 Oahu US 1/20/1963
186 Themeda villosa waipio valley PRESERVED_SPECIMEN o. degener & i. degener 29849 Oahu US 1/20/1963
187 Themeda villosa Honolulu PRESERVED_SPECIMEN Lee, H. A. 121 Oahu US 4/7/1905
188 Themeda villosa jct of hwy 18 and 19, above kailua-kona. PRESERVED_SPECIMEN d. r. herbst & s. ishikawa 5061 Hawaii US 10/7/1974
189 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii US 7/7/1997
190 Themeda villosa hilo forest reserve, 16mi marker on saddle rd PRESERVED_SPECIMEN c. r. annable, r. palmer, d. paul & d. ward 3604 Hawaii US 7/7/1997
191 Themeda villosa Hilo Forest Reserve, 16mi marker on Saddle Rd PRESERVED_SPECIMEN Annable, C. R.; Palmer, R.; Paul, D.; Ward, D. 3604 Hawaii US 7/7/1997
192 Themeda villosa Hanamaulu, Lihue. In pasture near beach. PRESERVED_SPECIMEN E. Y. Hosaka 1655 Kauai PTBG 11/4/1936
193 Themeda villosa kealakekua PRESERVED_SPECIMEN toy, b. collector number: s.n. Maui BISH 5/1/1979
194 Themeda villosa Jct of Hwy 18 and 19, above Kailua-Kona. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 5061 Hawaii US 10/7/1974
195 Themeda villosa south hilo distr., scenic drive along hamakua coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN staples, g.w. purell, m. collector number: 1268 Hawaii BISH 2/13/2007
196 Themeda villosa sw of dillingham ranch, mokuleia PRESERVED_SPECIMEN o. degener, d. l. topping, -- martinez & -- salucop 11058 Oahu US 3/12/1937
197 Themeda villosa Honolulu; Fair grounds PRESERVED_SPECIMEN Lee, H.A. 121 Oahu BISH 12/15/1924
198 Themeda villosa jeep road between wahiawa bog (kanaele swamp) and trail to tv relay tower on mt. kahili PRESERVED_SPECIMEN wagner, w.l. mill, s.; flynn,t.; peterson, b.; peterson, g. collector number: 5199 Kauai BISH 12/22/1983
199 Themeda villosa haleakala substation, makawao PRESERVED_SPECIMEN e. y. hosaka 2443 Maui US 4/9/1939
200 Themeda villosa SW of Dillingham Ranch, Mokuleia PRESERVED_SPECIMEN Degener, O.; Topping, D. L.; Martinez, --; Salucop, -- 11058 Oahu US 3/12/1937
201 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. Collector Number: 923 Hawaii BISH 11/2/1981
202 Themeda villosa hilo forest reserve; saddle rd. at 21.7 km ridge; with Holcus, Acacia, Hypochoeris, Styphelia PRESERVED_SPECIMEN annable, c.r. palmer, r. collector number: 3748 Maui BISH 1/14/1998
203 Themeda villosa hanamaulu, lihue, near beach PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
204 Themeda villosa Tree Planting Road near Stainback highway Ruderal. PRESERVED_SPECIMEN Stemmermann, R.L. 6703 Hawaii BISH 1/6/1981
205 Themeda villosa Hilo Forest Reserve; Saddle Rd. at 21.7 km PRESERVED_SPECIMEN Annable, C.R. 3748 Hawaii BISH 1/14/1998
206 Themeda villosa Kona; Keauhou PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 11/12/1937
207 Themeda villosa Kona Along roadside. PRESERVED_SPECIMEN Hosaka, E.Y. s.n. Hawaii BISH 9/1/1936
208 Themeda villosa Jeep road between Wahiawa Bog (Kanaele Swamp) and trail to TV relay tower on Mt. Kahili PRESERVED_SPECIMEN Wagner, W.L. 5199 Kauai BISH 12/22/1983
209 Themeda villosa Kawailoa Training Area, Drum Road, Kamananui Gulch Roadside vegetation, along with Hyparrhenia rufa, Dicranopteris linearis, Panicum maximum. PRESERVED_SPECIMEN Kawelo, K. s.n. Oahu BISH 6/4/2003
210 Themeda villosa koolau mountains; moanalua valley, damon estate Common along road up to parking area; associated with guava, koa, Schinus terebinthifolius, Digitaria pruriens. PRESERVED_SPECIMEN herbst, d.r. collector number: 355 Oahu BISH 3/18/1967
211 Themeda villosa Moanalua Valley, Campsite No. 1 Common along stream and jeep trail up to about 800 ft. PRESERVED_SPECIMEN Herbst, D.R. 1523 Oahu BISH 1/15/1970
212 Themeda villosa tree planting road near stainback highway Ruderal. PRESERVED_SPECIMEN stemmermann, r.l. collector number: 6703 Hawaii BISH 1/6/1981
213 Themeda villosa junction of highways 18 & 19, above kailua-kona PRESERVED_SPECIMEN herbst, d.r. ishikawa, s. collector number: 5061 Hawaii BISH 10/7/1974
214 Themeda villosa Laupahoehoe Section of Hilo Forest Reserve, North Hilo District Growing beside jeep road in logged 'ohi'a-koa forest. PRESERVED_SPECIMEN Cuddihy, L.W. 923 Hawaii BISH 11/2/1981
215 Themeda villosa honolulu; lyon arboretum, on makai side of access road ca. 200 yards from seismograph station. below section 31 Sunny, periodically sickled. PRESERVED_SPECIMEN baker, r.f. collector number: 61 Oahu BISH 4/9/1975
216 Themeda villosa hanamaulu, lihue, near beach In pasture. PRESERVED_SPECIMEN hosaka, e.y. collector number: 1655 Kauai BISH 11/4/1936
217 Themeda villosa South Hilo distr., Scenic Drive along Hamakua Coast, where forest ends and mac nut groves begin. Steep roadside bank at edge of mac nut plantation, in full sun on soil over lava substrate. PRESERVED_SPECIMEN Staples, G. 1268 Hawaii BISH 2/13/2007
218 Themeda villosa Honokohau Valley, Kava Kamp Roadside PRESERVED_SPECIMEN C. R. Annable 3748 Maui HAPI 1/14/1998
219 Themeda villosa Oahu Islands, Waianae Range, Waikele Gulch Disturbed roadside in wet lowland PRESERVED_SPECIMEN C. R. Annable 3604 Oahu HAPI 7/7/1997
220 Themeda villosa southwest of dillingham ranch Single plant in pasture. PRESERVED_SPECIMEN degener, o. topping, d.l.; martinez, m.; salucop, f. collector number: 11058 Oahu BISH 3/12/1937
221 Themeda villosa Southwest of Dillingham Ranch Single plant in pasture. PRESERVED_SPECIMEN Degener, O. 11058 Oahu BISH 3/12/1937
222 Themeda villosa kona PRESERVED_SPECIMEN hosaka, e.y. collector number: s.n. Oahu BISH 9/1/1936
223 Themeda villosa kona, before descent to kailua Wet area. PRESERVED_SPECIMEN christensen, c. collector number: 2 Hawaii BISH 11/14/1963
224 Themeda villosa koolau mountains; moanalua valley PRESERVED_SPECIMEN herbst, d.r. collector number: 355 BISH 3/18/1967
225 Themeda villosa Hoea PRESERVED_SPECIMEN C. R. Annable, R. Palmer, D. Paul & D. Ward 3604 Hawaii HAPI 7/7/1997